Our analyses reveal a single origin of the LT flock with the inclusion of the nonendemic S. Bayesian-relaxed clock analyses were used to estimate timings of radiations. Data were analysed using different methods of phylogenetic inference: Bayesian (also testing compositional heterogeneity), likelihood and parsi- mony, in order to investigate biogeographic history and the extent of intralacustrine speciation within this group. Here we estimate Synodontis relationships, based on mitochondrial and nuclear DNA, greatly expanding previous sampling. Synodontis catfish are a species-rich, tropical pan-African genus that predom- inately occur in fluviatile environments, but which also form a small radiation within Lake Tanganyika (LT).
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